Combine flours, wheat germ, brown sugar, baking powder, soda, salt, cinnamon, and nutmeg in a large bowl; make a well in center of mixture. Combine eggs, milk, and oil; add to dry ingredients, stirring just until moistened. Stir in apple and lemon rind.
Spoon into muffin pans coated with cooking spray, filling two-thirds full. Bake at 400F for 10 to 12 minutes.
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